Flowers have evolved an organ-specific type of metabolism where primary and specialized metabolites contribute to bolstering reproduction, the ultimate physiological function of flowers. Flower primary metabolites sustain development and the initial phases of fruit and seed set. Metabolites of the central pathway also serve as precursors for the synthesis of pigments, scents, and nectar, which attract and reward pollinators, as well as bioactive molecules that offer protection towards biotic and abiotic stresses. In the lab, we utilize a metabolomic approach to measure the whole spectrum of primary and specialized metabolites of flowers. We further integrate these measurements with transcriptomics and genomics and experiments with animal pollinators. Our research aims at understanding how metabolism changes during flower development and in response to climate change, and ultimately drives pollination in selfing and outcrossing species.